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The upper layers of mucus containing secreted mucins (e.g., MUC5AC, MUC5B) block larger debris while the mucins associated with proximity to the epithelium, known as membrane-tethered mucins (e.g., MUC1, MUC4, MUC16), block access to the cilia and its underlying epithelium for debris down to 40 nm in size (Kesimer et al., 2013)