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TOX/TOX2 and NR4A13 function as central regulators that cooperate to establish and maintain an exhaustion program through chromatin remodeling and epigenetic stabilization (90, 91)

The review further suggested that melanocortin receptor activation [may modulate] cellular resilience through modulation of redox sensitive pathways and pigment associated sequestration processes. These hypotheses were presented as exploratory and largely inferential, relying on mechanistic overlap rather than direct experimental validation

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