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In the nucleus, FOXO4 activates the transcription of genes encoding antioxidant enzymes like manganese superoxide dismutase (MnSOD), catalase (CAT), and glutathione peroxidase (GPX)

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Klys HS, Whillis D, Howard G, Harrison DJ (1992) Glutathione S-transferase expression in the human testis and testicular germ cell neoplasia

Likewise, Western blotting results showed that ICAM1, TNF, and IL6 protein levels were increased in the HG group compared with the NG group, which were inhibited by HU treatment (Figures 5GJ)